Ionis Pharmaceuticals Inc (IONS)

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Institutional Investors' Holdings

Filing Period:

No.	Name	Shares	Value	% Port
1.	Fmr Llc	24,072,101	$1,904,343,910	0.1%
2.	Capital World Investors	17,961,299	$1,420,918,364	0.19%
3.	Vanguard Group Inc	16,440,459	$1,300,604,712	0.02%
4.	Blackrock, Inc.	10,647,863	$842,352,443	0.01%
5.	AmundiBOND	3,200,000	$508,256,000	0.14%